Zoology

Zoology is the branch of biology devoted to the study of the animal kingdom (Animalia). This article discusses the history and concerns of that study. For a discussion of animals and a description of animal groups.

The current study of zoology has two main focuses: on particular taxonomic groups, and on the structures and processes common to most of them.

Taxonomically oriented studies concentrate on the different divisions of animal life. Invertebrate zoology deals with multicellular animals without backbones; its subdivisions include entomology (the study of insects) and malacology (the study of mollusks). Vertebrate zoology, the study of animals with backbones, is divided into ichthyology (fish), herpetology (amphibians and reptiles), ornithology (birds), and mammalogy (mammals). Paleontology, the study of fossils, is subdivided by taxonomic groups. In each of these fields, researchers investigate the classification, distribution, life cycle, and evolutionary history of the particular animal or group of animals under study. Most zoologists are also specialists in one or more of the process-oriented disciplines described below.

Morphology, the study of structure, includes gross morphology, which examines entire structures or systems, such as muscles or bones; histology, which examines body tissues; and cytology, which focuses on cells and their components. Many great advances made in cytology in recent years are attributable to the electron microscope and the scanning electron microscope. Special staining techniques and radioactive tracers have been used to differentiate structural detail at the molecular level. Methods have been developed for mapping neural connections between parts of the brain and for stimulating and recording impulses from specific brain sites and even individual nerve cells.

Anatomy

Anatomy (Greek anatomē, “dissection”), branch of natural science dealing with the structural organization of living things. It is an old science, having its beginnings in prehistoric times. For centuries anatomical knowledge consisted largely of observations of dissected plants and animals. The proper understanding of structure, however, implies a knowledge of function in the living organism. Anatomy is therefore almost inseparable from physiology, which is sometimes called functional anatomy. As one of the basic life sciences, anatomy is closely related to medicine and to other branches of biology.

It is convenient to subdivide the study of anatomy in several different ways. One classification is based on the type of organisms studied, the major subdivisions being plant anatomy (see Plant) and animal anatomy. Animal anatomy is further subdivided into human anatomy and comparative anatomy, which seeks out similarities and differences among animal types (see Animal). Anatomy can also be subdivided into biological processes—for example, developmental anatomy, the study of embryos, and pathological anatomy, the study of diseased organs. Other subdivisions, such as surgical anatomy and anatomical art, are based on the relationship of anatomy to other branches of activity under the general heading of applied anatomy. Still another way to subdivide anatomy is by the techniques employed—for example, microanatomy, which concerns itself with observations made with the help of the microscope (see the section below on the history of anatomy).

Related topics:

HUMAN ANATOMY
• Musculoskeletal System
• Nervous System
• Circulatory System
• Immune System
• Respiratory System
• Digestive and Excretory Systems
• The Endocrine System
• The Reproductive System

COMPARATIVE ANATOMY

Botany

Botany, branch of biology concerned with the study of plants (kingdom Plantae; see Plant). Plants are now defined as multicellular organisms that carry out photosynthesis. Organisms that had previously been called plants, however, such as bacteria, algae, and fungi, continue to be the province of botany, because of their historical connection with the discipline and their many similarities to true plants, and because of the practicality of not fragmenting the study of organisms into too many separate fields.

Botany is concerned with all aspects of the study of plants, from the smallest and simplest forms to the largest and most complex, from the study of all aspects of an individual plant to the complex interactions of all the different members of a complicated botanical community of plants with their environment and with animals (see Ecology).

Botany today does not depend on the fossil record for information concerning evolution and classification as much as does zoology, because the record for plants is much less complete than that for animals. Nevertheless, paleobotany, the study of fossil plants, has contributed greatly to the overall understanding of the evolution of the major groups of plants and especially to understanding of the interrelationships among the classes of seed plants. But much remains to be learned before fundamental questions such as the origin of the flowering plants (Angiosperm) can be answered.

Botanists—those engaged in the study of plants—occupy themselves with a broad range of activities. Many botanists are in academic positions that involve both teaching and research duties. The latter may involve laboratory work or field studies. Strictly speaking, botany is a pure science concerned with investigating the basic nature of plants. Many aspects of botany, however, have direct importance to human welfare and advancement, and applied botany is an important field. Such fields as forestry and horticulture are closely tied to basic botanical studies, whereas those such as pharmacology and agronomy are not as closely related but still depend on basic botanical knowledge.

Biology

Biology, the science of life. The term was introduced in Germany in 1800 and popularized by the French naturalist Jean-Baptiste de Lamarck as a means of encompassing the growing number of disciplines involved with the study of living forms. The unifying concept of biology received its greatest stimulus from the English zoologist Thomas Henry Huxley, who was also an important educator. Huxley insisted that the conventional segregation of zoology and botany was intellectually meaningless and that all living things should be studied in an integrated way. Huxley’s approach to the study of biology is even more cogent today, because scientists now realize that many lower organisms are neither plants nor animals (see Prokaryote; Protista). The limits of the science, however, have always been difficult to determine, and as the scope of biology has shifted over the years, its subject areas have been changed and reorganized. Today biology is subdivided into hierarchies based on the molecule, the cell, the organism, and the population.

Molecular biology, which spans biophysics and biochemistry, has made the most fundamental contributions to modern biology. Much is now known about the structure and action of nucleic acids and protein, the key molecules of all living matter. The discovery of the mechanism of heredity was a major breakthrough in modern science. Another important advance was in understanding how molecules conduct metabolism, that is, how they process the energy needed to sustain life.

Cellular biology is closely linked with molecular biology. To understand the functions of the cell—the basic structural unit of living matter—cell biologists study its components on the molecular level. Organismal biology, in turn, is related to cellular biology, because the life functions of multicellular organisms are governed by the activities and interactions of their cellular components. The study of organisms includes their growth and development (developmental biology) and how they function (physiology). Particularly important are investigations of the brain and nervous system (neurophysiology) and animal behavior (ethology).

Population biology became firmly established as a major subdivision of biological studies in the 1970s. Central to this field is evolutionary biology, in which the contributions of Charles Darwin have been fully appreciated after a long period of neglect. Population genetics, the study of gene changes in populations, and ecology, the study of populations in their natural habitats, have been established subject areas since the 1930s. These two fields were combined in the 1960s to form a rapidly developing new discipline often called, simply, population biology. Closely associated is a new development in animal-behavior studies called sociobiology, which focuses on the genetic contribution to social interactions among animal populations.

Biology also includes the study of humans at the molecular, cellular, and organismal levels. If the focus of investigation is the application of biological knowledge to human health, the study is often termed biomedicine. Human populations are by convention not considered within the province of biology; instead, they are the subject of anthropology and the various social sciences. The boundaries and subdivisions of biology, however, are as fluid today as they have always been, and further shifts may be expected.

See Animal; Animal Behavior; Botany; Cell; Classification; Development; Ecology; Evolution; Genetics; Heredity; Life; Medicine; Metabolism; Plant; Reproduction; Respiration; Zoology.

Ecology

Ecology, the study of the relationship of plants and animals to their physical and biological environment. The physical environment includes light and heat or solar radiation, moisture, wind, oxygen, carbon dioxide, nutrients in soil, water, and atmosphere. The biological environment includes organisms of the same kind as well as other plants and animals.

Because of the diverse approaches required to study organisms in their environment, ecology draws upon such fields as climatology, hydrology, oceanography, physics, chemistry, geology, and soil analysis. To study the relationships between organisms, ecology also involves such disparate sciences as animal behavior, taxonomy, physiology, and mathematics.

An increased public awareness of environmental problems has made ecology a common but often misused word. It is confused with environmental programs and environmental science (see Environment). Although the field is a distinct scientific discipline, ecology does indeed contribute to the study and understanding of environmental problems.

The term ecology was introduced by the German biologist Ernst Heinrich Haeckel in 1866; it is derived from the Greek oikos (“household”), sharing the same root word as economics. Thus, the term implies the study of the economy of nature. Modern ecology, in part, began with Charles Darwin. In developing his theory of evolution, Darwin stressed the adaptation of organisms to their environment through natural selection. Also making important contributions were plant geographers, such as Alexander von Humboldt, who were deeply interested in the “how” and “why” of vegetational distribution around the world.